This is the issue, the feed is bought once a year based on the animals at that time NOT when you look in the habitat. It is a bit simplistic but i do believe that is how it works. Also to the OP, you have a LOT of lions and elephants which are expensive animals. Money in this game is NOT realistic, it is designed to fit the gameplay.
On my third very large zoo now and this is the first I have managed to make profitable late game and the key to it was guest happiness / distribution of services. Happy guests, happy zoo.
on that point, the guest have not enough money, but dont leave the zoo because they are too happy. They still stay, where the are and are blocking the shops and no new guest with fresh money will enter. the solution: free the lions and let eat them the guest=> you dont need to feed them and the poor will leave (we should apply that in real life)
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